New Permian tetrapod footprints and macroflora from Turkey (Cakraz Formation, northwestern Anatolia): Biostratigraphic and palaeoenvironmental implications

Gand G., Tuysuz O., STEYER J. S., ALLAIN R., SAKINC M., SANCHEZ S., ...More

COMPTES RENDUS PALEVOL, vol.10, no.8, pp.617-625, 2011 (SCI-Expanded) identifier identifier

  • Publication Type: Article / Article
  • Volume: 10 Issue: 8
  • Publication Date: 2011
  • Doi Number: 10.1016/j.crpv.2011.09.002
  • Journal Indexes: Science Citation Index Expanded (SCI-EXPANDED), Scopus
  • Page Numbers: pp.617-625
  • Istanbul Technical University Affiliated: Yes


New tetrapod footprints belonging to the ichnogenus Hyloidichnus have been discovered in Turkey for the first time, in the lower part of the Cakraz Formation (Northwestern Anatolia) and together with macrofloral imprints of Annularia and Stigmaria. These discoveries confirm the Permian age of the fossiliferous red beds in which the coniferophyte Walchia was previously recorded. Based on the stratigraphic range of Annularia,Stigmaria and Hyloidichnus known elsewhere, a Cisuralian age is proposed for these beds. These new ichno- and macrofloral remains, together with the sedimentological data (mudcracks, rain drops) suggest the presence of captorhinid reptiles living in a palustrine floodplain environment, and under a warm, seasonal climate alternating between humid and relatively long dry seasons. These climatic conditions may have permitted the migration of these captorhinids through Laurasia during the Permian. (C) 2011 Academie des sciences. Published by Elsevier Masson SAS. All rights reserved.